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All pigment on the body, with the optional exception of those markings with Ghosting, has been removed from a colonist. You can often see the skin colour through the fur, usually pink or grey but sometimes affected by Artifice or a diluted version of the base coat colour. These colonists usually have red or pink eyes.
This rare trait causes dark markings to "double up" upon one another, merging into significantly thicker forms. On markings like spots or stripes, this can cause these shapes to grow and even appear layered upon each other. On markings like saddle or blanket, it can increase the range.